This was a bug in the code of MDEV-12387 "Push conditions into materialized
subqueries". The bug manifested itself in rather rare situations. An
affected query must contain IN subquery predicate whose left operand
was an outer field of a mergeable derived table or view and right operand
was a materialized subquery.
The erroneous code in fact stripped off the Item_direct_ref wrapper from
the left operand of the IN subquery predicate when building equalities
produced by the conversion of the predicate into a semi-join. As a result
the left operand was not considered as an outer reference anymore and
used_tables() was calculated incorrectly. This caused a crash in the
function optimize_keyuse().